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"ability to close quarters combat" - his shapeshifting obviously helps him because of his physical tools, but all of his progress to close quarters combat is from learning immortal techniques either from his manager or his dad.
"his intuition for fighting" - This one is a little muddy. Many of his fighting instincts are shown to be picked up from his mother beating him up a lot, but similar to the above he also picks up a ton of these from his manager.
"his ability to change his senses" - This was noted as something advanced immortals could do. This isn't a shapeshifting only trait.
Now going to your last comment, all of those things have been mentioned for shapeshifting; however, outside of the mom's training element, almost all of his progress since then has been because he has developed his immortal abilities not his shapeshifting abilities. This is what I mean when I'm complaining about them not playing into his shapeshifting more. Right now he has a passive shapeshifting buff from the very beginning of the story when he trained with his Mom's friend. Since then he hasn't developed anything on that side of the bloodline.