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You, like most humans believe it's related to DNA and the make-up of the brain, and it's perfectly reasonable from a scientific view. I don't. It's all about consciousness, not the other way around. So when his consciousness was put into muscle guy's body, all the knowledge and memories he had came with him too, which is why he can still use and understand magic principles.
You can argue that the hardware, the brain itself, doesn't run as well as his previous one, but that's bs because muscle guy has a much stronger and healthier body.
Most people play 1 time, and for dark souls-like games which are meant for replayability 2-3 is usually max so I don't understand what's your reasoning for saying that
Keep in mind we're talking about RPG games which can have up to dozens of hours of gameplay